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A Tyranid Bio-Titan leads a Tyranid Swarm in an irresistible onslaught against Imperial forces

A Bio-Titan is a monstrously large Tyranid bioform intended to counter the greatest war engines deployed by the other intelligent species of the Milky Way Galaxy. Towering above the battlefields, leading the swarms of lesser Tyranid creatures, stand the vast forms of the Bio-Titans. These massive biomechanical war-machines stalk the warzones of the Imperium spitting death and destruction, cutting down their foes with potent bio-weapons. Their approach is a terrifying sight to behold as they rip apart buildings, tear to pieces infantry and tank formations and even destroy Imperial Titans, Eldar Wraithlords or Ork Gargants with their armoured claws and ripper tentacles. Propelled relentlessly forward on their powerful legs, these horrifying creatures smash enemy defences into rubble with their deadly arsenal of bio-weapons. In return their enemies can do little; even under the heaviest fire, Bio-Titans are notoriously hard to kill as their huge bodies are protected by ridged plates of chitinous armour, and even when a lucky shot does penetrate their armoured sides a Bio-Titan is capable of rapid tissue regeneration.

Known Bio-Titan StrainsEdit

Carnifex/Dominatrix StrainEdit

Hierophant pict-capture

A massive Hierophant Tyranid Bio-Titan stalking the battlefield


A Tyranid Viciator, a Bio-Titan strain directly derived from the Hierophant

  • Hierophant - The Hierophant Bio-Titan is an immense, hideous creature, towering over the battlefields and bristling with spines, tendrils and symbiote weapons. They are amongst the largest biomorphs yet encountered by the forces of the Imperium amongst the swarms of the Hive Fleets, comparable in size and power to an Imperial Titan. A brood of Hierophants are likely to be at the forefront of a major Tyranid attack -- striding forwards, drawing enemy heavy weapons fire away from the lesser creatures of the swarm, absorbing huge amounts of damage before smashing into the enemy line with devastating force. The Hierophant is the Tyranids' largest shock trooper, armed with monstrous Bio-Cannons, multiple claws and razor-sharp blades, and armoured with a thick chitinous hide that exudes poisonous spores from rends in its plates. Like all Tyranid biomorphs, the Bio-Titan seems to be able to mutate its genome rapidly, evolving new organic weapons and defences. Other variants of the Hierophant Bio-Titan, such as the less well-known Viciator, have been identified with huge crushing claws or long scythe-like blades, cluster spines and other bio-weapons. How the Tyranids have developed such large biological constructs remains a mystery to the Imperium's Xenos Biologis. Whilst it seems a Hive Tyrant or Carnifex could easily be the result of a further genetic manipulation of the basic Tyranid Warrior genotype, there is no known genetic antecedent among the existing Tyranid bioforms for the Hierophant. It is speculated by the Magi Biologis of the Adeptus Mechanicus that the Hierophant was derived from the combined genetic strains of the Tyranid Carnifex and the Dominatrix. The forces of the Imperium have learned that combating Hierophants is simply a matter of applying the heaviest firepower available; large ordnance weapons and heavy artillery have so far proved the only effective weapons in destroying Bio-Titans.

Winged StrainEdit

Harridan Attack

A Tyranid Harridan attacks

It is speculated that the Harridan Bio-Titan is derived from a combined strain of the Gargoyle and the Winged Warrior subgroup of the Tyranid Warrior genus.

  • Harridan - The Harridan is the largest flying Tyranid bioform the Imperium has yet encountered in its bitter war against that inter-galactic alien menace. Harridans are truly massive creatures, likened to the flying drakes and wyverns of ancient Terran legend. They are the largest Tyranid bioform capable of atmospheric flight, soaring through a prey world’s skies on vast, leathery wings. Though they lack the sheer speed of an attack aircraft, it is a brave fool indeed who thinks them an easy target. They are the Gargoyles' "brood-mothers" and carry their "young" into battle. Circling high above, the Harridan waits patiently before diving down over the battlefield, strafing its foes with Bio-Cannons or swooping low to rake a tank with its two massive claws, or snapping its great fanged jaws around an infantryman. This creature's fearsome armament makes the Harridan capable of attacking just about any target on the field of battle. The Magos Biologis of the Adeptus Mechanicus have noted that Harridans have been encountered in a variety of sizes. Some are huge, upwards of 30 metres in length with a sizable wingspan of 40 metres, while other, far smaller Harridans have also been encountered that were well below the size of what would be considered a Bio-Titan. Instead, these Harridan variants are more like larger and somewhat tougher Gargoyles.

Other Bio-Titan StrainsEdit

Dactylis 2

The menacing Tyranid Dactylis

  • Dactylis - The Dactylis Bio-Titan is speculated to be a larger genetic derivative of the Tyranid Biovore. The Dactylis hurls large Spore Mines for very long ranges. Their forelimbs are long and covered in tightly-corded tendon and muscle that makes a distinctive cracking sound when used to hurl its deadly payload. When initially encountered by the forces of the Imperium, the Dactylis was noted for using the bio-weapon known as a Bile Pod. This living ammunition is a parasitic creature which feeds off the toxins emitted by the poisonous glands of the Dactylis' cranium. In recent encounters, the Dactylis has shown the Tyranids' innate ability to adapt to any combat situation or tactical need -- its cranial glands have grown a variety of spore-generating colonies that can be used to produce a number of different types of deadly ammunition that can be chosen for their particular utility against specific classes of foe.
Dominatrix conversion

A Tyranid Dominatrix

  • Dominatrix - The Dominatrix is a specialised Bio-Titan, commanding the forces on the battlefield and providing the highest level of psychic control as exceptionally powerful -- and valuable -- Tyranid synapse creatures. It is a fearsome opponent, for its sheer mass and panoply of bio-weaponry dwarf that of an Imperial super-heavy tank. The Dominatrix is no unthinking beast –- its level of intelligence is akin to that of the Hive Tyrant, and the strength of its telepathic link to the Hive Mind surpasses even those mighty creatures. Whilst within range of the Dominatrix's psychic aura, lesser Tyranid creatures become little more than an extension of the Dominatrix's will. As a result, a swarm under the Dominatrix's direct supervision functions with uncanny efficiency and a tactical genius that is beyond compare. Fortunately, such beasts are extremely rare in all but the most advanced Tyranid swarm, for where the Dominatrix treads, doom surely follows. Dominatrixes are the Tyranids' largest known Bio-Titans, and it is rumoured that they sometimes carry the Norn-Queens themselves into battle on their backs. They are armed with an immensely potent Warp blast weapon, Bio-plasma emitters, and their massive chitinous claws. They also serve as synapse creatures, the Hive Mind's primary battlefield coordinators for the Tyranid swarms. Because of this role, the destruction of a Dominatrix can severely incapacitate a Tyranid swarm that it is responsible for directing, though the firepower required to bring down a Dominatrix is immense.
Norn Queen Concept 2

A Magos Biologis sketch of a Tyranid Norn-Queen

  • Norn-Queen - Tyranid Norn-Queens fill the role of the "queen" of a Hive Fleet, similar to that of a Terran queen ant, termite, wasp or bee. As far as it is known, they are bound to a Tyranid Hive Ship and cannot leave it save under the most extreme of circumstances. They are responsible for the engineering and birthing of new Tyranids of every known bioform and they psychically communicate with the other Tyranid synapse creatures, directly transmitting the will of the Hive Mind. A Norn-Queen is always the most protected Tyranid aboard a Hive Ship, so the humans of the Imperium have been unable to garner much data about them save in the rare instances where they have been able to penetrate to the heart of a Tyranid swarm in deep space. The death of a Norn-Queen will sever the synaptic link between the Hive Mind and those Tyranids who are directly linked to that Norn-Queen, rendering them mindless, enraged beasts. The death of the Norn-Queen at Tarsis Ultra as a result of the heroic actions of Captain Uriel Ventris and the Ultramarines 4th Company caused the entire Tyranid swarm present on that world to turn on each other and become feral beasts. Norn-Queens also, to a great degree, control the evolutionary course of the Tyranid swarms. Upon arrival at a new world they intend to strip of its biomass, the Tyranids disgorge the "Alpha" swarm of biomorphs, which contain the genetic material of that particular fleet's Norn-Queen. These creatures then relay telepathic information to the Norn-Queen on the new world's environmental conditions and battlefield requirements, which spawn the second and subsequent generations of Tyranid bioforms, sometimes with tens of thousands of casualties during the collection of the initial data. Capture of an Alpha Wave Tyranid bioform can lead to the creation of a deadly neurotoxin that directly affects the Norn-Queen, as was done by the Ultramarines during the invasion of Ultramar by Hive Fleet Behemoth. When a Norn-Queen is killed, the psychic backlash of its death in the Hive Mind causes nearby Tyranid Hive Ships to calve, creating more Norn-Queens within their interiors. This is an instinctual survival mechanism known to the Magi Biologis of the Imperium as the Hydra Effect, and is the reason why even when a Hive Fleet is seemingly defeated, it can quickly form splinter fleets that in time become larger than the original.


  • Codex: Tyranids (6th Edition)
  • Codex: Tyranids (5th Edition)
  • Codex: Tyranids (4th Edition)
  • Codex: Tyranids (3rd Edition)
  • Imperial Armour Volume Four - The Anphelion Project
  • Warhammer 40,000 Collectible Card Game (Viciator Card)

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